The "servers are only on for a few hours" thing was like never true so I have no idea where that claim is coming from. The web performance test took more than a few hours to run alone and we had way more aggressive soaks for other workloads.
My recollection was that "write zeroes" just became a cheaper operation between '12 and '14.
A fun fact to distract from the awkwardness: a lot of the kernel work done in the early days was exceedingly scrappy. The port mapping stuff for memcached UDP before SO_REUSEPORT for example. FB binaries couldn't even run on vanilla linux a lot of the time. Over the next several years we put a TON of effort in getting as close to mainline as possible and now Meta is one of the biggest drivers of Linux development.